For enhanced safety, the front and second-row seat shoulder belts of the Land Rover Discovery have pretensioners to tighten the seatbelts and eliminate dangerous slack in the event of a collision and force limiters to limit the pressure the belts will exert on the passengers. The Toyota 4Runner doesn’t offer pretensioners for its second-row seat belts.
Both the Discovery and 4Runner have child safety locks to prevent children from opening the rear doors. The Discovery has power child safety locks, allowing the driver to activate and deactivate them from the driver's seat and to know when they're engaged. The 4Runner’s child locks have to be individually engaged at each rear door with a manual switch. The driver can’t know the status of the locks without opening the doors and checking them.
To provide maximum traction and stability on all roads, All-Wheel Drive is standard on the Discovery. But it costs extra on the 4Runner.
The Land Rover Discovery’s rear backup camera has a standard washer for maintaining a clear view under various conditions. In contrast, the Toyota 4Runner does not offer a rear camera washer, meaning its effectiveness relies on manual cleaning by the user when necessary.
To help make backing out of a parking space safer, the Discovery has standard Rear Traffic Monitor with Rear Traffic Braking, systems which detect vehicles approaching from the sides and can automatically apply the brakes to prevent a collision. Only the 4Runner Limited/Platinum/Trailhunter/TRD Pro offers Parking Support Brake.
For better protection of the passenger compartment, the Discovery uses safety cell construction with a three-dimensional high-strength frame that surrounds the passenger compartment. It provides extra impact protection and a sturdy mounting location for door hardware and side impact beams. The 4Runner uses a body-on-frame design, which has no frame members above the floor of the vehicle.
Both the Discovery and the 4Runner have standard driver and passenger frontal airbags, front side-impact airbags, side-impact head airbags, four-wheel antilock brakes, traction control, electronic stability systems to prevent skidding, crash mitigating brakes, daytime running lights, lane departure warning systems, blind spot warning systems, rearview cameras, rear cross-path warning and driver alert monitors.

